龙虱科
鞘翅目的一科昆虫
（重定向自龙虱）
此条目翻译品质不佳，原文在en:Wikipedia。 |
“龙虱”重定向至此。“龙虱”也可以指小型单人龙舟。
龙虱科（学名：Dytiscidae），亦作龙虱科，为水甲虫的一种，隶属于鞘翅目之下的肉食亚目。生活在田野、水沟、小溪等水体中，是一种掠食性水生昆虫。其学名源于希腊文δυτικός（dytikos），意思就是“肉食性的潜水甲虫”。本科物种几乎见于全世界所有淡水栖境，但亦有部分于落叶堆中栖息[1]。成虫体长一般在1和2.5 cm（0.4—1.0英寸）之间，尽管物种间的差异会比较大。当中见于欧洲的Dytiscus latissimus及巴西的Megadytes ducalis有最大的体型，可分别长至4.5 cm（1.8英寸）到4.75 cm（1.9英寸）[1][2]。与之相反，拥有最小体型的物种应当为栖息于澳大利亚地下水中的Limbodessus atypicali，只有约0.9 mm（0.035英寸）长[1]。 大多数物种的颜色都在深棕色、黑色或深橄榄绿，某些亚科的物种会在其边沿带有金黄色的条状斑纹。
龙虱科 化石时期：侏罗纪晚期至今
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_-_(imago),_Berkenwoude,_the_Netherlands.jpg)
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| Cybister lateralimarginalis | |
科学分类 
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| 界： | 动物界 Animalia |
| 门： | 节肢动物门 Arthropoda |
| 纲： | 昆虫纲 Insecta |
| 目： | 鞘翅目 Coleoptera |
| 亚目： | 肉食亚目 Adephaga |
| 总科： | 龙虱总科 Dytiscoidea |
| 科： | 龙虱科 Dytiscidae Leach, 1815 |
| 模式属 | |
| 龙虱属 Dytiscus Linnaeus，1758
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| 亚科、族与属 | |
| 多样性 | |
| 至少160个属，约4000物种 | |
食用
编辑本科物种的成虫，特别是Cybister属的物种，均为可食用昆虫。在美国内华达州洪堡干湖的一个山洞内，还在当地史前人类遗下的粪化石内发现有Cybister explanatus被食用的残骸[3]。在墨西哥，他们在吃C. explanatus时会以墨西哥夹饼夹着来吃。在日本多个地区（例如：长野县）有以日本龙虱（Cybister japonicus）为食的习惯；在中华人民共和国广东省，除了日本龙虱以外，亦会食用C. bengalensis、C. guerini、C. limbatus、C. sugillatus、C. tripunctatus等物种。D. marginalis很可能也包括在内，但这个物种的肉很少。也有报告指在台湾、泰国及新几内亚亦有食用本科物种的习惯[4]。
大但在陆地上很慢，成年后也不是特别凶猛，它们也被许多中型鸟类、哺乳动物和其他大型捕食者津津有味地吃掉。幼虫通常比较安全，由于它们的保护色和通过水柱逃跑的能力；它们可能相当难抓，可能成为小池塘的顶级掠食者。
系统分类
编辑- Acilius Leach, 1817
- Aethionectes Sharp, 1882
- Africodytes Biström, 1988
- Africophilus Guignot, 1948
- Agabetes Crotch, 1873
- Agabinus Crotch, 1873
- 豆龙虱属 Agabus Leach, 1817[6]，端毛龙虱属
- Agametrus Sharp, 1881
- Agaporomorphus Zimmermann, 1921
- Aglymbus Sharp, 1880
- Agnoshydrus Biström, Nilsson & Wewalka, 1997
- Allodessus Guignot, 1953
- 彩斑龙虱属 Allopachria Zimmermann, 1924
- Amarodytes Régimbart, 1900
- Amurodytes Fery & Petrov, 2013
- Andex Sharp, 1882
- Andonectes Guéorguiev, 1971
- Anginopachria Wewalka, Balke & Hendrich, 2001
- Anodocheilus Babington, 1841
- Antiporus Sharp, 1882
- Australphilus Watts, 1978
- Austrelatus Shaverdo, Hájek, Hendrich, Surbakti, Panjaitan & Balke, 2023
- Austrodytes Watts, 1978
- Barrethydrus
- Barretthydrus Lea, 1927
- Barretthydrus geminatus Lea, 1927
- Barretthydrus stepheni Watts, 1978
- Barretthydrus tibialis Lea, 1927
- Batrachomatus Clark, 1863
- Belladessus K.B.Miller & Short, 2015
- Bidessini Sharp, 1880
- Bidessodes Régimbart, 1895
- Bidessonotus Régimbart, 1895
- Bidessus Sharp, 1882
- Boreonectes (Fery & Ribera, 2018) Fery & Ribera, 2018
- Boreonectes Angus, 2010
- Borneodessus Balke, Hendrich, Mazzoldi & Biström, 2002
- Brachyvatus Zimmermann, 1919
- Brancuporus Hendrich, Toussaint & Balke, 2014
- Bunites Spangler, 1972
- Calicovatellus K.B.Miller & Lubkin, 2001
- Canthyporus Zimmermann, 1919
- Capelatus Turner & Bilton, 2015
- Caperhantus Balke, Hájek & Hendrich, 2017
- Carabdytes Balke, Hendrich & Wewalka, 1992
- Carabhudrus
- Carabhydrus Watts, 1978
- Carrhydrus
- Celina Aubé, 1837
- Chostonectes Sharp, 1880
- Clarkhydrus Fery & Ribera, 2018
- Clemnius Villastrigo, Ribera, Manuel, Millán & Fery, 2017
- Clypeodytes Régimbart, 1894
- Coelambus Thomson, 1860
- Coelambus cribrarius (Scudder, 1900)
- Coelambus derelictus (Scudder, 1900)
- Coelambus disjectus (Scudder, 1900)
- Coelambus dzieduszyckii (Lomnicki, 1894)
- Coelambus ganglbaueri (Lomnicki, 1894)
- Coelambus infernalis (Scudder, 1900)
- Coelambus latefasciatus (Lomnicki, 1894)
- Coelambus miocenus (Wickham, 1912)
- Coelambus niedzwiedzkii (Lomnicki, 1894)
- Coelambus ozokeriticus (Lomnicki, 1894)
- Coelambus picipoides (Lomnicki, 1894)
- Coelhydrus Sharp, 1882
- Colymbetes Clairville, 1806
- Colymbetes Schellenberg, 1806
- Comaldessus Spangler & Barr, 1995
- Copelatus Erichson, 1832
- Copelatus abonnenci Guignot, 1939
- Copelatus acamas Guignot, 1955
- Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017
- Copelatus advena Sharp, 1882
- Copelatus aemulus Bilardo & Rocchi, 1995
- Copelatus aequatorius Régimbart, 1899
- Copelatus aethiopicus Régimbart, 1906
- Copelatus agaroensis Balke, 2008
- Copelatus agrias Guignot, 1954
- Copelatus alternatus Sharp, 1882
- Copelatus amaroides Guignot, 1952
- Copelatus amatolensis Omer-Cooper, 1965
- Copelatus amazonicus Régimbart, 1889
- Copelatus ambiguus Bertrand & Legros, 1975
- Copelatus amphibius Ranarilalatiana & Bergsten, 2019
- Copelatus anastomosans Guignot, 1952
- Copelatus andobonicus Guignot, 1960
- Copelatus andreinii Régimbart, 1905
- Copelatus andrewesi J.Balfour-Browne, 1939
- Copelatus angolensis Peschet, 1924
- Copelatus angouei Bilardo & Rocchi, 2011
- Copelatus angustus Gschwendtner, 1932
- Copelatus ankaratra Ranarilalatiana & Bergsten, 2019
- Copelatus annobomensis J.Balfour-Browne, 1939
- Copelatus anthracinus Régimbart, 1895
- Copelatus antoniorum Hájek & Brancucci, 2011
- Copelatus aphroditae Balke, 2003
- Copelatus apicinotatus Bilardo & Rocchi, 2018
- Copelatus assalyi Bilardo & Rocchi, 2010
- Copelatus assimilis Régimbart, 1895
- Copelatus ateles Guignot, 1955
- Copelatus atrosulcatus Régimbart, 1906
- Copelatus babyrousa Hájek, Hendrich & Balke, 2018
- Copelatus bacchusi Wewalka, 1981
- Copelatus bacillifer Guignot, 1955
- Copelatus baculiformis Guignot, 1955
- Copelatus badeni Sharp, 1882
- Copelatus bakewelli J.Balfour-Browne, 1939
- Copelatus baoulicus Bilardo & Pederzani, 1978
- Copelatus bapounouensis Bilardo & Rocchi, 2019
- Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus barbouri Young, 1942
- Copelatus basifasciatus Régimbart, 1895
- Copelatus basilewskyi Bilardo & Pederzani, 1979
- Copelatus bechynei Guignot, 1953
- Copelatus befasicus Guignot, 1956
- Copelatus bengalensis Guignot, 1955
- Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996
- Copelatus betampona Ranarilalatiana & Bergsten, 2019
- Copelatus bezdeki Sheth, Ghate & Hájek, 2018
- Copelatus bibulus Guignot, 1948
- Copelatus bicolor (Fabricius, 1792)
- Copelatus biformis Sharp, 1882
- Copelatus bimaculatus (Resende & Vanin, 1991)
- Copelatus binaghii Bilardo & Pederzani, 1978
- Copelatus biolleyi Guignot, 1952
- Copelatus biroi Guignot, 1956
- Copelatus birougouensis Bilardo & Rocchi, 2018
- Copelatus biseriatus J.Balfour-Browne, 1939
- Copelatus biswasi Mukherjee & Sengupta, 1986
- Copelatus blancasi Guignot, 1958
- Copelatus blatchleyi Young, 1953
- Copelatus bolivianus Guignot, 1957
- Copelatus bombycinus Guignot, 1952
- Copelatus bonvouloiri Sharp, 1882
- Copelatus bottegoi Régimbart, 1895
- Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus boukali Hendrich & Balke, 1998
- Copelatus brancuccii Rocchi, 1979
- Copelatus brasiliensis Zimmermann, 1921
- Copelatus brendelli Hájek, Hendrich & Balke, 2018
- Copelatus brevicornis (Sharp, 1882)
- Copelatus brevistrigatus Guignot, 1959
- Copelatus brivioi Rocchi, 1976
- Copelatus bromeliarum (Scott, 1912)
- Copelatus brullei Aubé, 1838
- Copelatus brunneus J.Balfour-Browne, 1939
- Copelatus buqueti Aubé, 1838
- Copelatus burgeoni Gschwendtner, 1930
- Copelatus caelatipennis Aubé, 1838
- Copelatus caelatipennis Chevrolat, 1863
- Copelatus caelatus Guignot, 1952
- Copelatus caffer J.Balfour-Browne, 1939
- Copelatus calaquei Bilardo & Rocchi, 2008
- Copelatus camerunensis Guignot, 1941
- Copelatus capensis Sharp, 1882
- Copelatus carayoni Legros, 1949
- Copelatus carinatus Sharp, 1882
- Copelatus celinoides Guignot, 1952
- Copelatus cessaima Caetano, Bená & Vanin, 2013
- Copelatus chevrolati Aubé, 1838
- Copelatus chibcha Guignot, 1952
- Copelatus chinensis Regimbart, 1899
- Copelatus chipiriricus Guignot, 1957
- Copelatus chloroticus Régimbart, 1899
- Copelatus cinnamomeus Régimbart, 1895
- Copelatus clarki Sharp, 1882
- Copelatus collarti Gschwendtner, 1932
- Copelatus compertus Guignot, 1956
- Copelatus concii Bilardo, 1982
- Copelatus concolor Sharp, 1882
- Copelatus concolorans J.Balfour-Browne, 1939
- Copelatus confinis Bilardo & Rocchi, 1999
- Copelatus congo Gschwendtner, 1938
- Copelatus consimilis Bilardo & Rocchi, 2002
- Copelatus consors Sharp, 1882
- Copelatus cooperae Bilardo & Pederzani, 1972
- Copelatus cordovai Megna & Epler, 2012
- Copelatus cordylinoides J.Balfour-Browne, 1938
- Copelatus coxalis Sharp, 1882
- Copelatus crassus Régimbart, 1895
- Copelatus cryptarchoides Régimbart, 1899
- Copelatus cubaensis Schaeffer, 1908
- Copelatus curtistriatus Bilardo & Rocchi, 1995
- Copelatus curvispinis Bilardo & Rocchi, 2008
- Copelatus cynthiae Bilardo & Rocchi, 2008
- Copelatus daemeli Sharp, 1882
- Copelatus danyi Megna & Epler, 2012
- Copelatus darlingtoni Young, 1942
- Copelatus davidi Wewalka, 2017
- Copelatus debilis Sharp, 1882
- Copelatus deccanensis Sheth, Ghate & Hájek, 2018
- Copelatus decellei Bilardo, 1982
- Copelatus decemsulcatus Régimbart, 1895
- Copelatus deceptor Bilardo & Rocchi, 1995
- Copelatus dentatipenis Jiang, Hajek & Jia, 2022
- Copelatus depressus Sharp, 1882
- Copelatus descarpentriesi Bertrand & Legros, 1975
- Copelatus diffisus Guignot, 1939
- Copelatus dimorphus Sharp, 1882
- Copelatus distinctus Aubé, 1838
- Copelatus distinguendus Régimbart, 1903
- Copelatus diversistriatus Jiang, Hajek & Jia, 2022
- Copelatus djiboutensis Wewalka & Jäch, 2017
- Copelatus dolosus Guignot, 1956
- Copelatus doriae Sharp, 1882
- Copelatus doudouensis Bilardo & Rocchi, 2015
- Copelatus duodecimstriatus Aubé, 1838
- Copelatus duponti Aubé, 1838
- Copelatus efoutensis Bilardo & Rocchi, 1995
- Copelatus ejactus Omer-Cooper, 1965
- Copelatus ellai Bilardo & Rocchi, 1995
- Copelatus elutus Guignot, 1958
- Copelatus enganensis Guignot, 1940
- Copelatus epactus Guignot, 1948
- Copelatus erichsonii Guérin-Méneville, 1847
- Copelatus espinhasso Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7]
- Copelatus esteriensis Bilardo & Pederzani, 1978
- Copelatus eucritus Guignot, 1952
- Copelatus evanidus Bilardo & Rocchi, 1995
- Copelatus exaratus Sharp, 1882
- Copelatus externus Kirsch, 1873
- Copelatus falciformis Pederzani & Schizzerotto, 2017
- Copelatus fasciatus Bilardo & Rocchi, 1995
- Copelatus fastidiosus Guignot, 1959
- Copelatus feae Régimbart, 1888
- Copelatus felicis Hajek, Jiang & Jia, 2022
- Copelatus fernandensis Guignot, 1952
- Copelatus ferruginicollis Régimbart, 1895
- Copelatus ferus Guignot, 1953
- Copelatus festae Régimbart, 1899
- Copelatus fidschiensis Zimmermann, 1928
- Copelatus fijiensis Guignot, 1955
- Copelatus filiformis Sharp, 1882
- Copelatus fizpaci Bilardo & Rocchi, 2013
- Copelatus flavicans Guignot, 1952
- Copelatus flavidus Régimbart, 1895
- Copelatus florae Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7]
- Copelatus fluviaticus Guignot, 1957
- Copelatus fontanus J.Balfour-Browne, 1950
- Copelatus fossilis Říha, 1974
- Copelatus fractistriatus Bilardo & Rocchi, 1995
- Copelatus freudei Guignot, 1955
- Copelatus fryi J.Balfour-Browne, 1939
- Copelatus fulviceps J.Balfour-Browne, 1938
- Copelatus fuscipennis Sharp, 1882
- Copelatus fuscomaculatus Guignot, 1952
- Copelatus gabonicus Bilardo & Pederzani, 1978
- Copelatus galapagoensis G.R.Waterhouse, 1845
- Copelatus garambanus Guignot, 1955
- Copelatus gardineri Scott, 1912
- Copelatus geayi Régimbart, 1904
- Copelatus geniculatus Sharp, 1882
- Copelatus gentilis Sharp, 1882
- Copelatus gestroi (Sharp, 1882)
- Copelatus gigas Bilardo & Rocchi, 2019
- Copelatus glyphicus (Say, 1823)
- Copelatus griffinii Régimbart, 1899
- Copelatus gschwendtneri Guignot, 1939
- Copelatus guadelupensis Legros, 1948
- Copelatus guerini Aubé, 1838
- Copelatus guineensis Guignot, 1955
- Copelatus haemorrhoidalis Régimbart, 1883
- Copelatus hararensis Guignot, 1952
- Copelatus hardenbergi J.Balfour-Browne, 1950
- Copelatus hebeter Guignot, 1953
- Copelatus heterogynus Régimbart, 1899
- Copelatus hydroporoides (Murray, 1859)
- Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000
- Copelatus iguelaensis Bilardo & Rocchi, 2002
- Copelatus ilybioides Régimbart, 1895
- Copelatus imitator Bilardo & Rocchi, 2002
- Copelatus inaequalis Sharp, 1882
- Copelatus incognitus Sharp, 1882
- Copelatus indicus Sharp, 1882
- Copelatus inopinatus Bilardo & Rocchi, 1995
- Copelatus inornatus Sharp, 1882
- Copelatus insidiosus Bilardo & Rocchi, 1995
- Copelatus insolitus Chevrolat, 1863
- Copelatus instabilis Régimbart, 1897
- Copelatus instriatus Zimmermann, 1921
- Copelatus insuetus Guignot, 1941
- Copelatus insulanus Guignot, 1939
- Copelatus integer Sharp, 1882
- Copelatus internus Régimbart, 1904
- Copelatus interstriatus J.Balfour-Browne, 1939
- Copelatus inuber Guignot, 1952
- Copelatus ipiformis Régimbart, 1895
- Copelatus irinus Régimbart, 1899
- Copelatus irregularis W.J.MacLeay, 1871
- Copelatus ischius Guignot, 1956
- Copelatus jactator Guignot, 1955
- Copelatus jaechi Bilardo & Rocchi, 2016
- Copelatus jamaicensis Young, 1942
- Copelatus japonicus Sharp, 1884
- Copelatus jarrigei Legros, 1954
- Copelatus javanus Régimbart, 1883
- Copelatus jocosus Guignot, 1955
- Copelatus johannis Guignot, 1954
- Copelatus kalaharii Gschwendtner, 1935
- Copelatus kammuriensis Tamu & Tsukamoto, 1955
- Copelatus kaszabi Guignot, 1956
- Copelatus kely Ranarilalatiana & Bergsten, 2019
- Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus kilimandjarensis Bilardo & Pederzani, 1972
- Copelatus kindianus Guignot, 1954
- Copelatus kongouensis Bilardo & Rocchi, 1999
- Copelatus koreanus Mori, 1932
- Copelatus laccophilinus Sharp, 1882
- Copelatus laeticulus Sharp, 1882
- Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus lamottei Legros, 1954
- Copelatus lanzai Bilardo & Rocchi, 1995
- Copelatus laraensis Bilardo & Rocchi, 1995
- Copelatus lasckonyi Bilardo & Rocchi, 1995
- Copelatus latens Guignot, 1952
- Copelatus laticollis Régimbart, 1899
- Copelatus latifasciatus Bilardo & Rocchi, 1999
- Copelatus latipes Sharp, 1882
- Copelatus latus J.Balfour-Browne, 1950
- Copelatus leonardii Bilardo & Rocchi, 1999
- Copelatus leonensis Legros, 1954
- Copelatus lepersonneae Gschwendtner, 1943
- Copelatus lignosus Guignot, 1955
- Copelatus lineatipennis Guignot, 1955
- Copelatus lineatus (Guérin-Méneville, 1838)
- Copelatus longicornis Sharp, 1882
- Copelatus lootensi Guignot, 1955
- Copelatus louayensis Bilardo & Rocchi, 2004
- Copelatus luctuosus Guignot, 1939
- Copelatus luridescens Régimbart, 1889
- Copelatus luteocinctus Guignot, 1955
- Copelatus luteomaculatus Guignot, 1956
- Copelatus luzonicus Guignot, 1952
- Copelatus macellus Guignot, 1950
- Copelatus madoni Guignot, 1952
- Copelatus mahajanga Pederzani & Hájek, 2005
- Copelatus mahleri Holmen & Vazirani, 1990
- Copelatus makokouensis Bilardo & Rocchi, 1995
- Copelatus malaisei Guignot, 1954
- Copelatus malavergnorum Manuel & Ramahandrison, 2022
- Copelatus mancus Sharp, 1887
- Copelatus marginatus Sharp, 1882
- Copelatus marginipennis (Laporte, 1835)
- Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019
- Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018
- Copelatus masculinus Régimbart, 1899
- Copelatus massaicus Guignot, 1941
- Copelatus mathani Guignot, 1952
- Copelatus maushomi Sheth, Ghate & Hájek, 2018
- Copelatus mbokoensis Bilardo & Rocchi, 2006
- Copelatus melanogrammus Régimbart, 1883
- Copelatus mendax Bilardo & Rocchi, 2011
- Copelatus miaowangi Hajek, Jiang & Jia, 2022
- Copelatus minimus Bilardo & Rocchi, 1995
- Copelatus minor Bilardo & Pederzani, 1978
- Copelatus minutissimus J.Balfour-Browne, 1939
- Copelatus mocquerysi Régimbart, 1895
- Copelatus mohelicus Guignot, 1959
- Copelatus monticola Guignot, 1951
- Copelatus montivagus Young, 1942
- Copelatus mopanshanensis Jiang, Zhao & Hajek, 2022
- Copelatus mulangensis Bameul, 2003
- Copelatus mundus Sharp, 1882
- Copelatus mutabilis Bilardo & Rocchi, 1999
- Copelatus mvoungensis Bilardo & Rocchi, 2004
- Copelatus mysorensis Vazirani, 1970
- Copelatus nakamurai Guéorguiev, 1970
- Copelatus nangaensis Guignot, 1952
- Copelatus nanlingensis Hajek, Jiang & Jia, 2022
- Copelatus nauclerus Guignot, 1939
- Copelatus neavei J.Balfour-Browne, 1950
- Copelatus neelumae Vazirani, 1973
- Copelatus neogestroi Balke, 2008
- Copelatus neoguineensis Zimmermann, 1919
- Copelatus nguemai Bilardo & Rocchi, 2011
- Copelatus nigricans Sharp, 1882
- Copelatus nigricollis Paulino de Oliveira, 1880
- Copelatus nigrolineatus Sharp, 1882
- Copelatus nigropennis Zimmermann, 1927
- Copelatus nigrostriatus Régimbart, 1895
- Copelatus nilssoni Wewalka, 1982
- Copelatus nimbaensis Legros, 1958
- Copelatus nitens Bilardo & Rocchi, 1999
- Copelatus nitidus Sharp, 1882
- Copelatus nodieri Régimbart, 1895
- Copelatus normalis Erichson, 1847
- Copelatus notius Omer-Cooper, 1965
- Copelatus nzei Bilardo & Rocchi, 1999
- Copelatus oblitus Sharp, 1882
- Copelatus obscurus Sharp, 1882
- Copelatus occultus Bilardo & Rocchi, 1995
- Copelatus ogasawarensis Kamiya, 1932
- Copelatus onorei Pederzani & Rocchi, 1982
- Copelatus ornatipennis Zimmermann, 1926
- Copelatus owas Régimbart, 1895
- Copelatus pachys Guignot, 1955
- Copelatus pallidus Régimbart, 1895
- Copelatus paludorum Guignot, 1952
- Copelatus pandanorum Scott, 1912
- Copelatus panguana Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7]
- Copelatus pantosi Guignot, 1958
- Copelatus papuensis J.Balfour-Browne, 1939
- Copelatus parabaptus Guignot, 1955
- Copelatus parallelipipedus Régimbart, 1895
- Copelatus parallelus Zimmermann, 1920
- Copelatus pardii Rocchi, 1990
- Copelatus parisii Guignot, 1959
- Copelatus parumstriatus Gschwendtner, 1934
- Copelatus paryphes Guignot, 1955
- Copelatus pederzanii Bilardo & Rocchi, 1995
- Copelatus pereirai Guignot, 1955
- Copelatus peridinus Guignot, 1955
- Copelatus pinnifer Guignot, 1952
- Copelatus piriensis Omer-Cooper, 1965
- Copelatus platynotus Régimbart, 1895
- Copelatus polystrigus Sharp, 1882
- Copelatus ponomarenkoi Říha, 1974
- Copelatus portior Guignot, 1956
- Copelatus posticatus (Fabricius, 1801)
- Copelatus poungai Bilardo & Rocchi, 2006
- Copelatus predaveterus Miller, 2003
- Copelatus prolixus Sharp, 1882
- Copelatus prolongatus Sharp, 1882
- Copelatus propino Guignot, 1960
- Copelatus propinquus Régimbart, 1895
- Copelatus proximus Sharp, 1882
- Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019
- Copelatus pulchellus (Klug, 1834)
- Copelatus pulicarius Régimbart, 1895
- Copelatus pumilus Régimbart, 1895
- Copelatus punctatus Bilardo & Rocchi, 1995
- Copelatus punctulatus Aubé, 1838
- Copelatus puzhelongi Jiang, Hajek & Jia, 2022
- Copelatus quadrisignatus Régimbart, 1877
- Copelatus quadristriatus Pederzani & Schizzerotto, 2017
- Copelatus racenisi Guignot, 1951
- Copelatus ragazzii Régimbart, 1887
- Copelatus regimbarti Branden, 1885
- Copelatus restrictus Sharp, 1882
- Copelatus rimosus Guignot, 1952
- Copelatus rivalis Guignot, 1952
- Copelatus rocchii Bilardo, 1982
- Copelatus romani Zimmermann, 1924
- Copelatus rosulae Hajek, Jiang & Jia, 2022
- Copelatus royi Legros, 1958
- Copelatus rubiginosus Guignot, 1954
- Copelatus ruficapillus Régimbart, 1895
- Copelatus ruteri Legros, 1954
- Copelatus saegeri Guignot, 1955
- Copelatus safiotra Ranarilalatiana & Bergsten, 2019
- Copelatus sahlbergi J.Balfour-Browne, 1939
- Copelatus sallaei Sharp, 1882
- Copelatus sanfilippoi Bilardo & Pederzani, 1972
- Copelatus saverii Pederzani & Schizzerotto, 2017
- Copelatus scalptus Guignot, 1954
- Copelatus scaphites Guignot, 1955
- Copelatus schaefferi Young, 1942
- Copelatus schereri Wewalka, 1981
- Copelatus schuhi Hendrich & Balke, 1998
- Copelatus scutatus Guignot, 1952
- Copelatus scytalotus Guignot, 1956
- Copelatus senegalensis Lagar & Hernando, 1991
- Copelatus sexstriatus Sharp, 1882
- Copelatus sharpi Branden, 1885
- Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010
- Copelatus silvestrii Régimbart, 1903
- Copelatus simoni Régimbart, 1889
- Copelatus singularis Bilardo & Rocchi, 1995
- Copelatus sociennus J.Balfour-Browne, 1952
- Copelatus solitarius Sharp, 1882
- Copelatus sordidipennis Régimbart, 1895
- Copelatus speciosus Régimbart, 1892
- Copelatus spoliatus Guignot, 1955
- Copelatus stavropolitanus Říha, 1974
- Copelatus stillicidicola Bilardo & Rocchi, 1995
- Copelatus striatellus Boheman, 1848
- Copelatus striaticollis Lucas, 1857
- Copelatus striatopterus Aubé, 1838
- Copelatus striatulus Aubé, 1838
- Copelatus strigipennis (Laporte, 1835)
- Copelatus strigosulus (Fairmaire, 1878)
- Copelatus strinatii Guignot, 1958
- Copelatus striolatus Peschet, 1917
- Copelatus stygis Guignot, 1958
- Copelatus subdeficiens Régimbart, 1902
- Copelatus subsimilis Guignot, 1958
- Copelatus substriatus Kirsch, 1873
- Copelatus subterraneus Guéorguiev, 1978
- Copelatus sudrei Bameul, 2003
- Copelatus sulcatus Sharp, 1882
- Copelatus sulcipennis (Laporte, 1835)
- Copelatus sumbawensis Régimbart, 1899
- Copelatus suppar Guignot, 1956
- Copelatus supplementaris Régimbart, 1895
- Copelatus sylvaticoides Bilardo & Rocchi, 2013
- Copelatus sylvaticus Guignot, 1952
- Copelatus takakurai Satô, 1985
- Copelatus tanaus Guignot, 1953
- Copelatus taprobanicus Wewalka & Vazirani, 1985
- Copelatus tenebrosis Regimbart, 1880
- Copelatus tenebrosus Régimbart, 1880
- Copelatus tengchongensis Hajek, Jiang & Jia, 2022
- Copelatus teranishii Kamiya, 1938
- Copelatus terminalis Sharp, 1882
- Copelatus ternatensis Régimbart, 1899
- Copelatus terzanii Bilardo & Saverio, 2016
- Copelatus thiriouxi Peschet, 1917
- Copelatus thrasys Guignot, 1952
- Copelatus tibialis Sharp, 1882
- Copelatus tinctor Guignot, 1954
- Copelatus togoensis Régimbart, 1895
- Copelatus tomokunii Satô, 1985
- Copelatus tondangoyei Bilardo & Rocchi, 2013
- Copelatus tostus J.Balfour-Browne, 1950
- Copelatus trifilis Guignot, 1958
- Copelatus triglyphus Guignot, 1955
- Copelatus trilobatus Régimbart, 1895
- Copelatus tschaga Bilardo & Pederzani, 1972
- Copelatus tucuchiensis J.Balfour-Browne, 1939
- Copelatus tulagicus Guignot, 1942
- Copelatus uludanuensis Hendrich & Balke, 1995
- Copelatus undecimstriatus Aubé, 1838
- Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus usagarensis Zimmermann, 1926
- Copelatus vagatus Guignot, 1952
- Copelatus vagestriatus Zimmermann, 1919
- Copelatus vagus Bilardo & Rocchi, 1995
- Copelatus validus Sharp, 1882
- Copelatus vancli Hájek, Hendrich & Balke, 2018
- Copelatus vanninii Bilardo & Rocchi, 1999
- Copelatus variegatus Régimbart, 1895
- Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus venustus Bilardo & Rocchi, 1995
- Copelatus vigintistriatus Fairmaire, 1869
- Copelatus vigintisulcatus Régimbart, 1895
- Copelatus villiersi Guignot, 1950
- Copelatus virungaensis Bilardo, 1982
- Copelatus vitraci Legros, 1948
- Copelatus vivax Guignot, 1953
- Copelatus vokoka Ranarilalatiana & Bergsten, 2019
- Copelatus wallacei J.Balfour-Browne, 1939
- Copelatus waltoni J.Balfour-Browne, 1950
- Copelatus wayanadensis Manivannan & Madani, 2011
- Copelatus wewalkai Holmen & Vazirani, 1990
- Copelatus weymarni J.Balfour-Browne, 1947
- Copelatus xanthocephalus Régimbart, 1899
- Copelatus xiniudong Jiang, Hajek & Jia, 2022
- Copelatus yacumensis Guignot, 1957
- Copelatus yaguarete Scheers, 2018
- Copelatus yombensis Bilardo & Rocchi, 2015
- Copelatus zadiensis Bilardo & Rocchi, 1995
- Copelatus zanabato Manuel & Ramahandrison, 2022
- Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019
- Copelatus zimmermanni Gschwendtner, 1934
- Copelatus Sheth, Ghate & Hájek, 2018
- Coptotomus Say, 1830
- Cretodytes Ponomarenko, 1977
- Crinodessus K.B.Miller, 1997
- 大龙虱属 Cybister Curtis, 1827，真龙虱属
- Darwinhydrus Sharp, 1882
- Deronectes Sharp, 1882
- Derovatellus Sharp, 1882
- Desmopachria Babington, 1841
- Deuteronectes Guignot, 1945
- Deuteronectus
- Dimitshydrus Uéno, 1996
- Dyschirioides
- Dyticus Geoffroy, 1762
- Dytiscus Linnaeus, 1758
- Electruphilus Balker, Toledo & Hendrich, 2019
- Ereboporus K.B.Miller, Gibson & Alarie, 2009
- Eretes Laporte, 1833
- Etruscodytes Mazza, Cianferoni & Rocchi, 2013
- Exocelina Broun, 1886
- Fontidessus K.B.Miller & Spangler, 2008
- Geodessus Brancucci, 1979
- Gibbidessus Watts, 1978
- Glareadessus Wewalka & Biström, 1998
- Graphoderus Dejean, 1833
- Graptodytes Seidlitz, 1887
- Haideoporus Young & Longley, 1976
- Hamadiana Benetti, Short & Michat, 2019
- Hemibidessus Zimmermann, 1921
- Heroceras Guignot, 1950
- Heterhydrus Fairmaire, 1869
- Heterosternuta Strand, 1935
- Hintonia Young, 1980
- Hoperius Fall, 1927
- Hornectes Fery & Ribera, 2018
- Hovahydrus Biström, 1982
- Huxelhydrus Sharp, 1882
- Hydaticus Leach, 1817
- Hyderodes Hope, 1838
- Hydrocolus Roughley & Larson, 2000
- Hydrodessus J.Balfour-Browne, 1953
- Hydrodytes K.B.Miller, 2001
- 短褶龙虱属 Hydroglyphus Motschulsky, 1853
- Hydronebrius Jakovlev, 1897
- Hydropeplus Sharp, 1882
- Hydroporus Clairville, 1806
- Hydroporus Schellenberg, 1806
- Hydrotrupes Sharp, 1882
- Hydrovatus Motschulsky, 1853
- 水龙虱属 Hygrotus Stephens, 1828
- Hyphoporus Sharp, 1882
- Hyphovatus Wewalka & Biström, 1994
- 异爪龙虱属 Hyphydrus Illiger, 1802
- Hypodessus Guignot, 1939
- Iberonectes Fery & Ribera, 2018
- Iberoporus Castro & Delgado, 2001
- Ilybiosoma Crotch, 1873
- Ilybius Erichson, 1832
- Incomptodessus K.B.Miller & García, 2011
- Isonotus Houlbert, 1934
- Japanolaccophilus Satô, 1972
- Kakadudessus Hendrich & Balke, 2009
- Kakadudessus tomweiri Hendrich & Balke, 2009
- Kuschelydrus Ordish, 1976
- Laccodytes Régimbart, 1895
- Laccomimus Toledo & Michat, 2015
- Lacconectus Motschulsky, 1855
- 粒龙虱属 Laccophilus Leach, 1815
- Laccoporus J.Balfour-Browne, 1939
- Laccoporus nigritulus (Gschwendtner, 1936)
- Laccornellus Roughley & Wolfe, 1987
- Laccornis Gozis, 1914
- Laccosternus Brancucci, 1983
- Lancetes Sharp, 1882
- Laodytes Queney, Lemaire & Ferrand, 2020
- Larsonectes Fery & Ribera, 2018
- Leconectes Fery & Ribera, 2018
- Leiodytes Guignot, 1936
- Leuronectes Sharp, 1882
- Liadyxianus Prokin, Petrov, Wang & Ponomarenko, 2013
- Limbodessus Guignot, 1939
- Liodessus Guignot, 1939
- Lioporeus Guignot, 1950
- Liopterus Dejean, 1833
- Madaglymbus Shaverdo & Balke, 2008
- Matus Aubé, 1836
- Megadytes Sharp, 1882
- Megaporus Brinck, 1943
- Meladema Laporte, 1835
- Melanodytes Seidlitz, 1887
- Meridiorhantus Balke, Hájek & Hendrich, 2017
- Mesodytes Prokin, Petrov, Wang & Ponomarenko, 2013
- Metaporus Guignot, 1945
- Methles Sharp, 1882
- Microdessus Young, 1967
- Microdytes J.Balfour-Browne, 1946
- Microhydrodytes K.B.Miller, 2002
- Miodytiscus Wickham, 1911
- Morimotoa Uéno, 1957
- Mystonectes Fery & Ribera, 2018
- Napodytes Steiner, 1981
- Nartus Zaitzev, 1907
- 孔龙虱属 Nebrioporus Régimbart, 1906
- Necterosoma Sharp, 1882
- Necterosoma W.J.MacLeay, 1871
- Nectoboreus Fery & Ribera, 2018
- Nectomimus Fery & Ribera, 2018
- Nectoporus Guignot, 1950
- Neobidessodes Hendrich & Balke, 2009
- Neobidessus Young, 1967
- Neoclypeodytes Young, 1967
- Neonectes J.Balfour-Browne, 1944
- Neonectes Zimmermann, 1932
- Neoporus Guignot, 1931
- Neoscutopterus J.Balfour-Browne, 1943
- 彩裳粒龙虱属 Neptosternus Sharp, 1882
- Notaticus Zimmermann, 1928
- Novadessus K.B.Miller, 2016
- Onychohydrus Schaum & White, 1847
- Oreodytes Seidlitz, 1887
- Pachydrus Sharp, 1882
- Pachynectes Régimbart, 1903
- Palaeodytes Ponomarenko, 1987
- Papuadessus Balke, 2001
- Paroster Sharp, 1882
- Peschetius Guignot, 1942
- Petrodessus K.B.Miller, 2012
- Philaccolilus Guignot, 1937
- Philaccolus Guignot, 1937
- Philodytes J.Balfour-Browne, 1939
- Phreatodessus Ordish, 1976
- Platambus Thomson, 1859
- Platydytes Biström, 1988
- Platynectes Gueorguiev, 1972
- Platynectes Régimbart, 1879
- Porhydrus Guignot, 1945
- Primospes Sharp, 1882
- Procoelambus Théobald, 1937
- Prodaticus Sharp, 1882
- 缝龙虱属 Pseuduvarus Biström, 1988
- Psychopomporus Jean, Telles & K.B.Miller, 2012
- Pteroporus Guignot, 1933
- Queda Sharp, 1882
- Regimbartina Chatanay, 1911
- Rhantaticus Sharp, 1880
- Rhantus Dejean, 1833
- Rhithrodytes Bameul, 1989
- Rompindessus Balke, Bergsten & Hendrich, 2017
- Sandracottus Sharp, 1882
- Sanfilippodytes Franciscolo, 1979
- Scarodytes Gozis, 1914
- Schistomerus Palmer, 1957
- Sekaliporus Watts, 1997
- Sharphydrus Omer-Cooper, 1958
- Siamoporus Spangler, 1996
- Siettitia Abeille de Perrin, 1904
- Sinodytes Spangler, 1996
- Spanglerodessus K.B.Miller & García, 2011
- Spencerhydrus Sharp, 1882
- Sternhydrus Brinck, 1945
- Sternopriscus Sharp, 1880
- Stictonectes Brinck, 1943
- Stictotarsus Zimmermann, 1919
- Stygoporus Larson & LaBonte, 1994
- Suphisellus Zimmermann, 1921
- Tassilodytes Fery & Bouzid, 2016
- Tepuidessus Spangler, 1981
- Thermonectus Dejean, 1833
- Thermonectus alfredi Griffini, 1898
- Thermonectus basillaris (Harris, 1829)
- Thermonectus batesi Sharp, 1882
- Thermonectus circumscriptus (Latreille, 1809)
- Thermonectus cuneatus Sharp, 1882
- Thermonectus depictus Sharp, 1882
- Thermonectus duponti (Aubé, 1838)
- Thermonectus intermedius Crotch, 1873
- Thermonectus laporti (Aubé, 1838)
- Thermonectus leprieuri J.Balfour-Browne, 1944
- Thermonectus margineguttatus (Aubé, 1838)
- Thermonectus marmoratus (Gray, 1831)
- Thermonectus nigrofasciatus (Aubé, 1838)
- Thermonectus nobilis Zimmermann, 1924
- Thermonectus sibleyi Goodhue-McWilliams, 1981
- Thermonectus simulator Sharp, 1882
- Thermonectus succinctus (Aubé, 1838)
- Thermonectus tremouillesi Michat & Torres, 2016[8]
- Thermonectus variegatus (Dejean, 1836)
- Thermonectus variegatus (Laporte de Castelnau, 1835)
- Thermonectus zimmermani Goodhue-McWilliams, 1981
- Tikoloshanes Omer-Cooper, 1956
- Tiporus Watts, 1985
- Trichonectes Guignot, 1941
- Trogloguignotus Sanfilippo, 1958
- Tyndallhydrus Sharp, 1882
- Typhlodessus Brancucci, 1985
- Uvarus Guignot, 1939
- Vatellus Aubé, 1837
- Yola Gozis, 1886
- Yolina Guignot, 1936
- Zaitzevhydrus Fery & Ribera, 2018
- Zimpherus K.B.Miller & Wheeler, 2015
参考文献
编辑- ^ 1.0 1.1 1.2 G.N. Foster; D.T. Bilton. The Conservation of Predaceous Diving Beetles: Knowns, Unknowns and Anecdotes. D.A. Yee (编). Ecology, Systematics, and the Natural History of Predaceous Diving Beetles (Coleoptera: Dytiscidae). 2014: 437–462. ISBN 978-94-017-9109-0.
- ^ Archived copy. [2015-05-19]. （原始内容存档于2015-05-21）.
- ^ Miller, Kelly; Bergsten, Johannes. Diving Beetles of the World: Systematics and Biology of the Dytiscidae. Baltimore: Johns Hopkins University Press. 3 October 2016: 20.
- ^ De Foliart (2002), Jäch (2003), CSIRO (2004)
- ^ Dytiscidae Species List. Joel Hallan's Biology Catalog. Texas A&M University. [2012-05-07]. （原始内容存档于2012-10-23） （英语）.
- ^ 6.0 6.1 Dytiscidae 龍蝨科. TaiBIF. [2019-03-13]. （原始内容存档于2019-07-28） （中文（繁体））.
- ^ 7.0 7.1 7.2 Jiří Hájek, Yves Alarie, Cesar J. Benetti, Neusa Hamada, Monika Springer, Lars Hendrich, Adrián Villastrigo, Rodulfo Ospina Torres, Michael S. Basantes and Michael Balke. 2024. Underestimated Diversity and Range Size of Diving Beetles in Tank Bromeliads—Coleoptera of ‘hygrofloric’ lifestyle (Dytiscidae). Zoological Journal of the Linnean Society. 200(3); 720–735. DOI: 10.1093/zoolinnean/zlad093
- ^ Mariano C. Michat & Patricia L. M. Torres (2016) Thermonectus tremouillesi sp. nov. (Coleoptera: Dytiscidae: Aciliini): description of the adults and larvae and comparisons with other species of the genus, Journal of Natural History, 50:25-26, 1633-1648, DOI: 10.1080/00222933.2016.1145274
- 联邦科学与工业研究组织 (CSIRO) (2004): Water for a Healthy Country - Family Dytiscidae （页面存档备份，存于互联网档案馆）. Version of 2004-JUL-02. Retrieved 2008-AUG-04
- De Foliart, Gene R. (2002): Chapter 26 - Eastern Asia: China, Japan, and other countries. In: The Human Use of Insects as a Food Resource: A Bibliographic Account in Progress.
- Jäch, Manfred A. (2003): Fried water beetles Cantonese style. American Entomologist 49(1): 34-37. PDF fulltext
- Larson, D.J., Alarie, Y., and Roughley, R.E. (2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa. ISBN 978-0-660-17967-4.
